Even if it were possible to hold ones breath indefinitely, the authors note toward the end of the paper, another human breathing nearby, or several meters upwind, would create a CO2 plume that could lead mosquitoes close enough to you that they may lock on to your visual signature. Efforts to protect our skin from mosquitoes on a summer nigh may work for a while, but not forever. The temperature sensor is required to be in contact with the object physically and it uses conduction for monitoring the changes in temperature. Use heat detectors in areas where fumes or smoke from a fire would inhibit the visibility of the fire. But it also paints a bleak picture for those hoping to avoid mosquito bites. 3. Which sense is that? The scientists reasoned that if they put the human diphtheria toxin receptor under the control of genetic sequences that normally regulate different TRPs, they could selectively eliminate certain neurons with a simple toxin injection. We sense temperature in our environment through specialized nerve cells that project into the outer layers of the skin. Mice without either TRPV1- or TRPM8-expressing cells were indifferent to temperatures between 0 and 50C. Free nerve endings for temperature have ion channels that respond to particular temperatures, while other free nerve endings generate action potentials in response to extreme force on the skin or other potentially damaging stimuli that is felt as pain. Then, guided by visual cues that draw it even closer, the mosquito can sense the hosts body heat. (Scientists could only detect the difference by touching the surfaces.) There are two zones of attention: the real zone of close is very small. Heat detector can sense the heat (change in heat according to the features of heat detector used). Much of this information reaches us through the 5 senses organs. See more at www . We absorb information through sight , hearing , touch , smell and taste, which is quickly relayed and interpreted to our brains to provoke a . Thermal sensors can also read heat signatures inside buildings, potentially revealing sensitive details about what happens behind the closed door of a home or office. This quiz is all about using our senses to detect energy and to find out what is going on around us. To test this hypothesis, the researchers did the same CO2 plume experiment, but this time they provided a dark object on the floor of the wind tunnel. ","blurb":"","authors":[{"authorId":8948,"name":"Frank Amthor","slug":"frank-amthor","description":"

Frank Amthor, PhD, is a professor of psychology at the University of Alabama and holds a secondary appointment in the UAB Medical School Department of Neurobiology. Rate-of-rise heat detectors sense the heat in the room. To comply with the new e-Privacy directive, we need to ask for your consent - Published in the journal Current Biology, the work provides researchers with exciting new information about insect behavior and may even help companies design better mosquito traps in the future. {"appState":{"pageLoadApiCallsStatus":true},"articleState":{"article":{"headers":{"creationTime":"2016-09-11T22:12:53+00:00","modifiedTime":"2016-09-11T22:12:53+00:00","timestamp":"2022-09-14T18:15:52+00:00"},"data":{"breadcrumbs":[{"name":"Academics & The Arts","_links":{"self":"https://dummies-api.dummies.com/v2/categories/33662"},"slug":"academics-the-arts","categoryId":33662},{"name":"Science","_links":{"self":"https://dummies-api.dummies.com/v2/categories/33756"},"slug":"science","categoryId":33756},{"name":"Neuroscience","_links":{"self":"https://dummies-api.dummies.com/v2/categories/33767"},"slug":"neuroscience","categoryId":33767}],"title":"How the Skin Senses Temperature and Pain","strippedTitle":"how the skin senses temperature and pain","slug":"skin-senses-temperature-pain","canonicalUrl":"","seo":{"metaDescription":"You can detect more than just various kinds of pressure on your skin. After training, the dogs were tested on their skill in double-blind experiments; all three successfully detected the objects emitting weak thermal radiation, the scientists reveal today in Scientific Reports. Dogs' noses just got a bit more amazing. This sense combines with taste to produce flavors. Warmth receptors respond best to particular temperatures above body temperature (98.6 degrees Fahrenheit), while cold receptors respond best to particular temperatures below body temperature. We make sense of this information based on previous experience (and subsequent learning) and by the combination of the information from each of the senses. What do we use to sense smell? \"https://sb\" : \"http://b\") + \".scorecardresearch.com/beacon.js\";el.parentNode.insertBefore(s, el);})();\r\n","enabled":true},{"pages":["all"],"location":"footer","script":"\r\n

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This was a new finding for flies, and we suspected that mosquitoes would exhibit a similar behavior. It also has a high temperature range from -20 o to 80 o C. The key component of a thermal camera is a heat sensor attached to a special type of lens, which is then adapted to work alongside standard image-capture technologies. Five senses: the interface Attention: the shining torch Our attention is like a torch, shining into areas where we are interested. PIR sensors and infrared detectors are inexpensive. They can work out which part of the pit is hottest, and therefore the direction of the heat source, which could be a warm-blooded prey animal. This flow begins a signal thats relayed to the brain. How do we do that? Lasers illuminate objects in an environment and reflect the . Whether it's to pass that big test, qualify for that big promotion or even master that cooking technique; people who rely on dummies, rely on it to learn the critical skills and relevant information necessary for success. Another ion channel called TRPM8 was subsequently identified as a low-temperature sensor. His research is focused on retinal and central visual processing and neural prostheses. One of the first heat detectors was a sprinkler system used in the 1860s. For unlimited access to all quizzes, games and more, you'll need to subscribe. Those are the eyes, ears, nose, tongue, and skin. 2022 American Association for the Advancement of Science. SMELL. Writer, educator, and avid student of nature. April 15, 2015 at 9:00 a.m. EDT. "[It] provides yet another window into the sensory worlds of dogs' highly evolved cold noses.". When they sense it, they initiate a cascade of signals that ends up modulating the activity of the hormone auxin. Information collected from our five primary senses helps detect danger and allows our brains to understand our environmental conditions better, enabling us to react to our surroundings. The . Copyright 2016-2022 - Education Quizzes Sweet and salty tastes are often a sign of nutrient-rich foods. My new book is "The Stickler's Guide to Science in the Age of Misinformation". Eating with our ears sounds somewhat funny, but when we think about our five senses, we do use our ears when we eat: we listen for the frying of fish or the crunch of potato chips. Thermal imaging cameras are handheld electronic devices with an integrated visual display, designed for detecting heat energy. As with sight, auditory processing relies on how the brain interprets, recognises and differentiates sound stimuli. Ion channels such as TRPV1 are essentially pores in the cell membrane that control the flow of electrically charged ions into and out of cells. 2013 Mar 27;33(13):5533-41. doi: 10.1523/JNEUROSCI.5788-12.2013. Infrared sensors can be found everywhere in our daily lives. It's published bythe Office of Communications and Public Liaison in the NIH Office of the Director. What did you see? Ears. These receptors have similar structures, or, really, lack of structure. When a concentrated CO2 plume was present, the mosquitos followed it within the tunnel as expected. Additional senses & variations. This system is responsible for all the sensations we feel - cold, hot, smooth, rough, pressure, tickle, itch, pain, vibrations, and more. Sam is looking at the stars. Echolocation We move our body, turn our head, and in doing so we focus and limit what is available for our senses. They found that in the presence of the carbon dioxide plumes, the mosquitoes were attracted to the dark high-contrast object. The two basic physical types of temperature sensors are Contact Temperature Sensor Types - The contact temperature sensor can be used to detect liquids, solids or gases over a wide range. Linear Heat Detection (LHD) is a continuous heat detector designed to detect heat along the length of a sensor cable. A metal solder with a low melting point was used to keep caps in place in a network of water-filled pipes. We sense temperature in our environment through specialized nerve cells that project into the outer layers of the skin. Sensors are sophisticated devices that are frequently used to detect and respond to electrical or optical signals. That is, we predicted that when the mosquitoes were exposed to CO2, which is an indicator of a nearby host, they would also spend a lot of time hovering near high-contrast objects, such as a black object on a neutral background.. So you could think of it as a type of memory or lasting effect., Next, the researchers wanted to see how a mosquito factors thermal information into its flight path. Extreme temperatures cause Mrgprd neurons to join in and signal an alarm. Obviously, we know that if you have an object in the presence of a CO2 plumewarm or coldthey will fly toward it because they see it, he says. But these gadgets are far from original. Pain receptors also exist that respond to chemical damage from acids or bases, and other types of damage such as that caused by a cut. This detector used the melting of materials to detect heat. These experiments show that the attraction to a visual feature and the attraction to a warm object are separate. 1. Specifically this quiz is aimed at the section dealing with using our senses to detect energy. He has been an NIH-supported researcher for over 20 years and has published over 100 journal articles and conference abstracts.

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Frank Amthor is a professor of psychology at the University of Alabama at Birmingham, where he also holds secondary appointments in the UAB Medical School Department of Neurobiology, the School of Optometry, and the Department of Biomedical Engineering. . The independent and iterative nature of the sensory-motor reflexes renders mosquitoes host seeking strategy annoyingly robust.. When the researchers put the human diphtheria toxin receptor under the control of TRPV1 regulatory sequences and injected diphtheria toxin into adult mice, TRPV1-expressing neurons were destroyed. in high hazard areas, such as boiler rooms, where a fire would be difficult to see. It senses its preys heat and by using vibrations in the ground. How many of these questions about our senses can you answer? It has a resolution of better than 0.1C and works similar to that of Thermocouple Temperature Sensors. The final puff: Can New Zealand quit smoking for good? Pain receptors also exist that respond to chemical damage from acids or bases, and other types of damage such as that caused by a cut.\r\n\r\nPain and temperature sensations tend to be carried by small caliber axons in a spinal cord tract called the lateral spinothalamic tract. A more medial pathway called the lemnical tract carries fine touch information via larger caliber axons with faster conduction velocities. Information gathered from all of these experiments enabled the researchers to create a model of how the mosquito finds its host over different distances. They are typically called free nerve endings. Free nerve endings for temperature have ion channels that respond to particular temperatures, while other free nerve endings generate action potentials in response to extreme force on the skin or other potentially damaging stimuli that is felt as pain. There's an app for that. And a whole lot of senses that regulate your internal state. Thermal imaging has important government uses . So we had to find a way to separate the visual attraction from the thermal attraction.. Several other related TRP (transient receptor potential) channels were also found to be stimulated by heating or cooling. Use heat detectors for early detection of fires. When observed under a microscope, guard hairs from mice resemble optical sensors used in thermal cameras to detect heat, according to a new study. A bitter or sour taste was an indication of poisonous plants or of rotting protein-rich food. Warmth receptors respond best to particular temperatures above body temperature (98.6 degrees Fahrenheit), while cold receptors respond best to particular temperatures below body temperature. A monitored smoke is a smoke detector that is connected with an alarm system in some way. Warmth receptors respond best to particular temperatures above body temperature (98.6 degrees Fahrenheit), while cold receptors respond best to particular temperatures below body temperature. In the wind tunnel with no CO2 plume, the insects ignored the dark object entirely. But mammals are also able to detect more pleasant cool and warm temperatures. What do people wear so they can hear better? Through the use of all these senses, dogs can literally smell and taste their surroundings and build up a very complete picture. The ancient Greek philosopher and scientist, Aristotle (384-322 BC), first described the five senses of sight, hearing, touch, taste, and smell; we now know we have many more. ","description":"You can detect more than just various kinds of pressure on your skin. Dummies helps everyone be more knowledgeable and confident in applying what they know. Within the somatosensory system, there are four main types of receptors: mechanoreceptors, thermoreceptors, pain receptors, and proprioceptors. Multiband. Frank Amthor is a professor of psychology at the University of Alabama at Birmingham, where he also holds secondary appointments in the UAB Medical School Department of Neurobiology, the School of Optometry, and the Department of Biomedical Engineering. A heat detector is usually used in rooms where there is excessive dust or fumes - for instance, a garage or cellar - which a smoke alarm . Bethesda, MD 20892-2094, How timing of eating affects metabolism and weight gain, Long COVID symptoms linked to inflammation, Cyberbullying linked with suicidal thoughts and attempts in young adolescents, E-cigarettes linked with blood vessel damage. "The study is consistent with other research that describes the combined dog nose and brain as a sophisticated platform for processing a broad range of signals," says Gary Settles, an emeritus professor of mechanical engineering at Pennsylvania State University, University Park, who has studied dogs' sniffing abilities. Amanda Heidt Amanda Heidt Amanda is an associate editor at The Scientist, where she oversees the Scientist to Watch, Foundations, and Short Lit columns. Use heat detectors where there is a potential for a fire to smolder for some time before igniting. Receptors for temperature and pain look like the axon terminals without any other structure around them. Additional resources. Researchers from the University of Washington contributed to the work, which was funded by the National Institutes of Health. As humans, we are capable of sensing temperatures above or below our body temperature. Most mammals have naked, smooth skin on the tips of their noses around the nostrils, an area called the rhinarium. In rare cases, as in certain insect antennae, TRPA1 is used to detect heat. At hotter or colder temperatures, these neurons become more active. Biology Coordination and Control The Five Senses 1 Answer Charlie P. Jun 25, 2018 Sight Smell Taste Touch Hearing Explanation: Sight -important as it allows us to see our surroundings in visible light, helps us to navigate, hunt, feed. This suggests that while these neurons may be responsible for detecting temperature, further study will be needed to identify all the receptors involved. While it was no surprise to see the mosquitoes tracking a CO2 plume, the new part that we found is that the CO2 plume increases the likelihood that theyll fly toward an object. Extreme heat, cold or skin pressure, however, activates receptors that are interpreted as pain. What is he using? There are six types of optical detectors commonly used in the process industries: 1. Have you ever looked through a pair of binoculars? Their study appeared on March 27, 2013, in the Journal of Neuroscience. As it flies closerto within 5 to 15 meters (16 to 52 feet)it begins to see the host. Heat detectors are special devices that are designed to sense a rapid change in air temperature and set off alarms based on the amount of temperature rise above ambient condition, a fixed-temperature point and rate of temperature rise. An intuition is simply the powerful sense that something is true without having an awareness or understanding of the reasons behind this feeling it may or may not represent something true about the world. We can taste our food. Science and AAAS are working tirelessly to provide credible, evidence-based information on the latest scientific research and policy, with extensive free coverage of the pandemic. Sensation detecting nerves found in this system are called sensory nerves and are activated by different sensations, be it temperature, pain, or tactile sense (touch). The strong link between taste and emotions has to do with our evolution: Taste helped us "test" the food we ate, so it was important for our survival. Eliminating TRPV1 and TRPM8 neurons has a stronger effect than knocking out only these genes. Sharks have the same 5 senses as we do but can also sense electrical currents and pressure changes. Some receptors those having what are called transient receptor potential (TRP) channels respond to both.\r\n\r\n[caption id=\"attachment_225072\" align=\"aligncenter\" width=\"246\"] Free nerve ending receptors for temperature and pain. If we've learned anything from the COVID-19 pandemic, it's that we cannot wait for a crisis to respond. An example of a sensor used in some robots is called LIDAR (Light Detection And Ranging). Magnifying glasses make things look bigger. An infrared photo of a golden retriever in a functional magnetic resonance imaging scanner reveals the cold temperature of a dog's nose versus its glowing, warm body. Proprioception : This sense gives you the ability to tell where your body parts are, relative to other body parts. Bibliography. If you wish to subscribe straight away, visit our Join Us page. Next, the researchers scanned the brains of 13 pet dogs of various breeds in a functional magnetic resonance imaging scanner while presenting the pooches with objects emitting neutral or weak thermal radiation. Lots of older people have hearing aids. Pain and temperature sensations tend to be carried by small caliber axons in a spinal cord tract called the lateral spinothalamic tract. Marco is bird-watching. You judge a wide range of temperatures (cool, damp, chilly, cold, warm, humid, hot, and so on) by sensing the unique ratio of activation of the different receptors activated at any particular temperature.\r\n\r\nExtreme heat, cold or skin pressure, however, activates receptors that are interpreted as pain. A laser sensor is very precise in measurement and in the same time is very . But some animals are able to detect infrared waves, which radiate from warm objects. Tongue. Two other skin senses are temperature and pain. The sensor is intended as an alternative to pyroelectric sensors or PIR detectors in home automation, building automation, healthcare . Light, sound and heat are all types of energy. Tom wants to look closely at this butterfly. Smell. Have you ever looked through a telescope? The Flir C5 is great for general use but also perfect as a thermal camera to detect mice and pests. into a signal which can be measured electrically.Sensor can be defined as an element that senses in one form of energy to produce a variant in same or another form of energy . The white horizontal line at the bottom of Carter's front door shows where warm air is escaping. This allows engineers to quickly identify regions . Light, sound and heat are all types of energy. But only very small changes in TRPA1 protein structure are required to transform it into an odor detector instead of a heat detector. These five sense organs contain receptors that relay information through the sensory neurons to the appropriate places within the nervous system. Hearing. These mice actually preferred higher temperatures than control mice. But, a circuit is to be designed for activating an alarm system to indicate fire or heat change and for alerting the security or protection system. A sense is a biological system used by an organism for sensation, the process of gathering information about the world through the detection of stimuli. A more medial pathway called the lemnical tract carries fine touch information via larger caliber axons with faster conduction velocities. The ability to sense weak, radiating heat is known in only a handful of animals: black fire beetles, certain snakes, and one species of mammal, the common vampire bat, all of which use it to hunt prey. Up to two thirds of the total weight of a shark's brain is dedicated to smell. They are used in TVs and TV remotes, police radars, astronomy tools, and security equipment. This type of sensor is triggered by the "rate of change" rather than the temperature itself. It provides us with unconscious information of. The dogs weren't able to see or smell the difference between these objects. That includes venomous snakes from the subfamily Crotalinae, commonly known as pit vipersso-called for the pair of heat-sensing organs located in "pits" between their eyes and . Even at temperatures that you would think that the ion channels are not doing anything, theyre actually sending out a small signal to the brain, Hoon suggests. In control experiments, researchers introduced a plume consisting of background air with a low concentration of CO2. They are typically called free nerve endings. Free nerve endings for temperature have ion channels that respond to particular temperatures, while other free nerve endings generate action potentials in response to extreme force on the skin or other potentially damaging stimuli that is felt as pain. You judge a wide range of temperatures (cool, damp, chilly, cold, warm, humid, hot, and so on) by sensing the unique ratio of activation of the different receptors activated at any particular temperature.\r\n\r\nExtreme heat, cold or skin pressure, however, activates receptors that are interpreted as pain. References: J Neurosci. Only when the mosquito gets closer does it detect an objects thermal signature, van Breugel says. Find out more. Skin. TJS - Web Design Lincolnshire. The somatosensory system includes the following cutaneous or skin senses: pressure (touch), pain (nociception), vibration and temperature, position sense (proprioception), and body movement (kinesthesis). Help News from Science publish trustworthy, high-impact stories about research and the people who shape it. Add your information below to receive daily updates. Together, the two experiments show that dogs, like vampire bats, can sense weak hot spots and that a specific region of their brains is activated by this infrared radiation, the scientists say. ___ It's possible for a vivid experience of consciousness to exist undetected from the outside ___ And when we inspect our intuitions . The detectors register 70 degrees as a baseline; when the heat in the room rises rapidly above 70 degrees, the alarm is activated. This quiz addresses the requirements of the National Curriculum KS1 Science for children aged 5 and 6 in years 1 and 2. Ultraviolet (UV); 2. Smell : Yet another of the sensors that work off of a chemical reaction. NIH Research Matters When an adult female mosquito needs a blood meal to feed her young, she searches for a hostoften a human. Subscribe to News from Science for full access to breaking news and analysis on research and science policy. Or take a look around the website and start at our Home page. To do this, the researchers constructed two glass objects that were coated with a clear chemical substance that made it possible to heat them to any desired temperature. Some receptors those having what are called transient receptor potential (TRP) channels respond to both. These receptors have similar structures, o","noIndex":0,"noFollow":0},"content":"You can detect more than just various kinds of pressure on your skin. . They found that mosquitoes showed a preference for the warm object. Please make a tax-deductible gift today. Specifically this quiz is aimed at the section dealing with using our senses to detect energy. This causes cells on the shaded side of . A laser light is very useful for tracking and detection a target located at a long distance. A heat detector could provide a solution. The heat detector circuit can be designed using a heat sensor. However, mosquitoes can also pick up other cues that. Futurity is your source of research news from leading universities. The sense of space. Gut Microbes Affect Weight After Gastric Bypass, Research Hints at New Approach to Addiction, Subscribe to get NIH Research Matters by email, Mailing Address: They hypothesize that from 10 to 50 meters away, a mosquito smells a hosts CO2 plume. 2. Thermal image cameras ("infrared imaging") pick up the heat emitted by an object, so they excel at tracking people and animals. New sense discovered in dog noses: the ability to detect heat. What are the 5 senses used for in science? Ultraviolet/infraredsimple voting (UV/IR); 5. AAAS is a partner of HINARI, AGORA, OARE, CHORUS, CLOCKSS, CrossRef and COUNTER. The left somatosensory cortex in dogs' brains, which delivers inputs from the nose, was more responsive to the warm thermal stimulus than to the neutral one. Although different types of pain receptors work by different mechanisms, what they have in common is that the sense of pain signals impending damage to the skin. Thermosensation the ability to detect temperature triggers our reflex to withdraw from painful heat or cold. They have antennae that they use to detect a pheromone scent trail that's been left behind by another termite to notify others in their colony to a food source. We can see light, hear sound and feel heat. The somatosensory system reacts to diverse stimuli and skin sensations through the mechanical, chemical, and thermal properties of the tissue . Which food do you not like the taste of? Funding:NIH National Institute of Dental and Craniofacial Research (NIDCR). The sense of smell is used for various purposes - sexual attraction, recognizing the odor of their own species, finding suitable sites in which to lay eggs, and finding food. What do we mainly use to taste our food? Our experiments suggest that female mosquitoes do this in a rather elegant way when searching for food. Dr. Cho believes that sound is the forgotten flavor sense and may not get the credit it deserves when we think about sensory evaluation. Equilibrioception (vestibular sense): Refers to the sense of balance and orientation in the space around us. A research team led by Dr. Mark A. Hoon of NIHs National Institute of Dental and Craniofacial Research (NIDCR) devised a way to target different classes of neurons. Many insects, mosquitoes included, are attracted by the odor of the carbon dioxide (CO2) gas that humans and other animals naturally exhale. Thermosensation the ability to detect temperature triggers our reflex to withdraw from painful heat or cold. This helps ensure that they dont waste their time investigating false targets like rocks and vegetation. This new study used cutting edge cell ablation technology to delete the nerve circuit that encodes heat and some forms of itch while preserving the circuitry that sense cold temperatures.. Firefighting thermal imagers (TIs) bolster our senses and improve decision making capability because they detect heat energy, specifically long-wave infrared energy, and are largely unaffected by . Not all smells are horrid. Sometimes our hearing is not very good. Dual-frequency infrared (IR/IR); 4. But touch (skin) comes before sight, or even before speech. It is the first expression and the last, and it always mentions the truth. Such as those generated by the Earth's geomagnetic field . Your tax-deductible contribution plays a critical role in sustaining this effort. However, mosquitoes can also pick up other cues that signal a human is nearby: they use their vision to spot a host and thermal sensory information to detect body heat. In my recent column, Different States of Matter, I alluded to the fact that the teachers at my high school were somewhat sparing with the truth. How do we do that? It is difficult to test, van Breugel says. This sensor cable is made up of two conductors, both insulated by heat sensitive polymers. Which foods do you like the taste of? You can find more about this topic by visiting. With its 160 x 120 thermal sensor, you will be able to do pest detection fast and the sensor is also backed up by Flir's MSX image enhancement technology which combines the image from the built-in visible light camera together with the thermal layer. Original Study Mrgprd neurons were previously believed to be involved only in detecting painful mechanical stimuli. Engineers must create them, as sensors, for robots. (For example, in the human body, the brain which is part of the central nervous system receives signals from the senses which continuously receive information from the environment, interprets these signals, and causes the body to respond . By combining information from both pits, the snake can also estimate the distance of the object. Beneath this jacket and heat sensitive polymer coating is a . It's likely that other vampire bats also use similar sensory organs to . Some receptors those having what are called transient receptor potential (TRP) channels respond to both.\r\n\r\n[caption id=\"attachment_225072\" align=\"aligncenter\" width=\"246\"] Free nerve ending receptors for temperature and pain. Many insects, mosquitoes included, are attracted by the odor of the carbon dioxide (CO2) gas that humans and other animals naturally exhale. We can see light, hear sound and feel heat. As currently defined, taste sensations fall into four, or possibly five, categories: sweet, sour, salty, and bitter; many Japanese researchers also argue that the taste of monosodium glutamate (MSG), which they refer to as umami, is unique and "basic" (for example, Nakamura and Kurihara, 1991; Rogers and Blundell, 1990). "When you see this . In one set of experiments, a high-concentration CO2 plume was injected into the tunnel, mimicking the signal created by the breath of a human. He wants to see the birds clearly.

","authors":[{"authorId":8948,"name":"Frank Amthor","slug":"frank-amthor","description":"

Frank Amthor, PhD, is a professor of psychology at the University of Alabama and holds a secondary appointment in the UAB Medical School Department of Neurobiology. He has been an NIH-supported researcher for over 20 years and has published over 100 journal articles and conference abstracts.

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